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More on the "Big Five" Five-Factor Model of Personality

The study also examined correlations between each aspect, and gender differences were small: absolute values 0 to 0.12 with a median of 0.03 and a mean of 0.04, with correlations having a scale of -1 to 1 (perfect negative to perfect positive correlation).

The correlations were somewhat consistent with the plasticity-stability supertraits (P: E, O ... S: A, C, -N), and were consistent with a General Personality Factor (E, A, C, -N, O). Notice that Neuroticism is inverted, making Emotional Stability. It has negative correlations with the other factors, while those factors have positive correlations among themselves.

The factors' correlations I find to have a mean of 0.19 and a standard deviation of 0.05.
 
Goodbye to MBTI, the Fad That Won't Die | HuffPost - from 2013

"My name is Adam Grant, and I am an INTJ." Something he learned from a Myers-Briggs Type Indicator test. A few months after scoring as INTJ, he scored as ESFP -- the opposite. "I began to read through the evidence, and I found that the MBTI is about as useful as a polygraph for detecting lies."

As my inconsistent scores foreshadowed, the MBTI does poorly on reliability. Research shows “that as many as three-quarters of test takers achieve a different personality type when tested again,” writes Annie Murphy Paul in The Cult of Personality Testing, “and the sixteen distinctive types described by the Myers-Briggs have no scientific basis whatsoever.” In a recent article, Roman Krznaric adds that “if you retake the test after only a five-week gap, there’s around a 50 percent chance that you will fall into a different personality category.”

... Although there are data suggesting that different occupations attract people of different types, there is no convincing body of evidence that types affect job performance or team effectiveness. As management researchers William Gardner and Mark Martinko write in a comprehensive review, “Few consistent relationships between type and managerial effectiveness have been found.”
The article: Using the Myers-Briggs Type Indicator to Study Managers: A Literature Review and Research Agenda - William L. Gardner, Mark J. Martinko, 1996
 
Ways where the MBTI fails:
  • Exhibit A: in the MBTI, thinking and feeling are opposite poles of a continuum. In reality, they’re independent: we have three decades of evidence that if you like ideas and data, you can also like people and emotions. (In fact, more often than not, they go hand in hand: research shows that people with stronger thinking and reasoning skills are also better at recognizing, understanding, and managing emotions.) When I scored as a thinker one time and a feeler one time, it’s because I like both thinking and feeling. I should have separate scores for the two.
  • Exhibit B: the feeling type is supposed to tap into my orientation toward people and emotions. But this lumps together three separate traits that capture a positive orientation toward others, the tendency to feel negative emotions, and the receptivity toward these emotions.
Consistent with earlier research and evaluations, there was no support for the view that the MBTI measures truly dichotomous preferences or qualitatively distinct types; instead, the instrument measures four relatively independent dimensions. The interpretation of the Judging-Perceiving index was also called into question. The data suggest that Jung's theory is either incorrect or inadequately operationalized by the MBTI and cannot provide a sound basis for interpreting it. However, correlational analyses showed that the four MBTI indices did measure aspects of four of the five major dimensions of normal personality.
 
An ambivert is someone who is halfway between being an introvert and an extrovert. Introversion and extroversion are two extremes of a continuum with the middle being ambiversion and most people clustering around that state.
Why does the MBTI remain so popular in spite of these problems? Murphy Paul argues that people cling to the test for two major reasons. One is that thousands of people have invested time and money in becoming MBTI-certified trainers and coaches. As I wrote over the summer, it’s awfully hard to let go of our big commitments. The other is the “aha” moment that people experience when the test gives them insight about others — and especially themselves.
 
The article then got into the Big Five model of personality.

Personality and Performance at the Beginning of the New Millennium: What Do We Know and Where Do We Go Next? - Barrick - 2001 - International Journal of Selection and Assessment - Wiley Online Library
Results support the previous findings that conscientiousness is a valid predictor across performance measures in all occupations studied. Emotional stability was also found to be a generalizable predictor when overall work performance was the criterion, but its relationship to specific performance criteria and occupations was less consistent than was conscientiousness. Though the other three Big Five traits (extraversion, openness and agreeableness) did not predict overall work performance, they did predict success in specific occupations or relate to specific criteria.
C'ness predicts academic and career success, and Rubenzer & Faschingbauer find most Presidents high in c'ness.

Deep-level composition variables as predictors of team performance: A meta-analysis. - PsycNET
he strength of the team composition variable and team performance relationships was moderated by the study setting (lab or field) and the operationalization of the team composition variable. In lab settings, team minimum and maximum general mental ability and team mean emotional intelligence were related to team performance. Only negligible effects were observed in lab settings for the personality factor and team performance relationships, as well as the value and team performance relationships. In contrast, team minimum agreeableness and team mean conscientiousness, openness to experience, collectivism, and preference for teamwork emerged as strong predictors of team performance in field studies.
 
Personality Development: Stability and Change | Annual Review of Psychology

Testing predictions from personality neuroscience. Brain structure and the big five - PubMed
We used a new theory of the biological basis of the Big Five personality traits to generate hypotheses about the association of each trait with the volume of different brain regions. Controlling for age, sex, and whole-brain volume, results from structural magnetic resonance imaging of 116 healthy adults supported our hypotheses for four of the five traits: Extraversion, Neuroticism, Agreeableness, and Conscientiousness. Extraversion covaried with volume of medial orbitofrontal cortex, a brain region involved in processing reward information. Neuroticism covaried with volume of brain regions associated with threat, punishment, and negative affect. Agreeableness covaried with volume in regions that process information about the intentions and mental states of other individuals. Conscientiousness covaried with volume in lateral prefrontal cortex, a region involved in planning and the voluntary control of behavior. These findings support our biologically based, explanatory model of the Big Five and demonstrate the potential of personality neuroscience (i.e., the systematic study of individual differences in personality using neuroscience methods) as a discipline.
They didn't find anything that covaried with Openness to Experience.

Empirical, Theoretical, and Practical Advantages of the HEXACO Model of Personality Structure - Michael C. Ashton, Kibeom Lee, 2007
Has a sixth factor: Honesty-Humility

The HEXACO Personality Inventory - Revised
 
Big Five or Big Six?

The HEXACO Personality Inventory - Revised
Has four facets for each factor:
  • Honesty-Humility -- Sincerity, Fairness, Greed Avoidance, Modesty
  • Emotionality -- Fearfulness, Anxiety, Dependence, Sentimentality
  • eXtraversion -- Social Self-Esteem, Social Boldness, Sociability, Liveliness
  • Agreeableness -- Forgivingness, Gentleness, Flexibility, Patience
  • Conscientiousness -- Organization, Diligence, Perfectionism, Prudence
  • Openness -- Aesthetic Appreciation, Inquisitveness, Creativity, Unconventionality
Has an additional combined factor: Altruism

Honesty‐Humility, the Big Five, and the Five‐Factor Model - Ashton - 2005 - Journal of Personality - Wiley Online Library - In the Big Five model, some of H-H resides in Agreeableness.

On Measuring the Sixth Basic Personality Dimension: A Comparison Between HEXACO Honesty-Humility and Big Six Honesty-Propriety - Isabel Thielmann, Benjamin E. Hilbig, Ingo Zettler, Morten Moshagen, 2017 - assessing two possible versions of a sixth personality factor: Honesty-Humility and Honesty-Propriety. Of the two, H-H is better supported than H-P.

The Mini-IPIP6: Tiny yet highly stable markers of Big Six personality - ScienceDirect -- all six factors' scores are highly stable across retesting.

Comparative validity of Brief to Medium-Length Big Five and Big Six Personality Questionnaires. - PsycNET
" In this study, 3 popular brief to medium-length Big Five measures (NEO Five Factor Inventory, Big Five Inventory [BFI], and International Personality Item Pool), and 3 six-factor measures (HEXACO Personality Inventory, Questionnaire Big Six Scales, and a 6-factor version of the BFI) were placed in competition to best predict important student life outcomes."
with result
"Six-factor inventories demonstrated better predictive ability for life outcomes than did some Big Five inventories."
 
Recurrent Personality Dimensions in Inclusive Lexical Studies: Indications for a Big Six Structure - Saucier - 2009 - Journal of Personality - Wiley Online Library
with reprint
untitled - Saucier 2009 JP Big Six Structure.pdf

This study worked on words from 7 languages (Chinese, English, Filipino, Greek, Hebrew, Spanish, and Turkish), and not just English.

The Big Six:
  • Conscientiousness: Diligent, Precise (6); Conscientious, --Irresponsible (5); Disciplined, Orderly, --Lazy, --Reckless (4); Dutiful, Industrious, Meticulous, Organized, Thorough, --Absent-minded, --Careless, --Disorderly, --Frivolous, --Rash (3); Hard-working, Methodical, Scrupulous, Steadfast, --Chaotic, --Imprudent, --Inaccurate, --Inattentive, --Inconsiderate, --Inconstant, --Irrational, --Lax, --Negligent, --Undisciplined, --Untidy, --Wishy-washy (2).
  • Honesty/Humility: Honest (6), Sincere, --Hypocritical (5); Loyal, --Conceited, --Greedy (4); Just, --Boastful, --Calculating, --Dishonest, --Sly (3); Altruistic, Modest, Truthful, --Naughty, --Lying, --Pompous, --Pretending, --Pretentious, --Stingy, --Untruthful (2).
  • Agreeableness: Peaceful, Tolerant, --Aggressive, --Choleric (5); Mild, Patient (4); Agreeable, Good-natured, --Authoritarian, --Hot-headed, --Irritable, --Stubborn (3); Accommodating, Conciliatory, Kind-hearted, Lenient, Sympathetic, Tranquil, Warm, --Brusque, --Explosive, --Fierce, --Irascible, --Quarrelsome, --Quick-tempered, --Short-tempered (2).
  • Emotionality: Vulnerable (6); Emotional (5); Anxious, Sentimental, --Courageous, --Self-Assured, --Strong (4); Fragile, --Brave, --Imperturbable, --Independent, --Resolute (3); Delicate, Depressive, Fearful, Hypersensitive, Indecisive, Insecure, Melancholic, Oversensitive, Suggestible, Whining, Worrying, --Bold, --Intrepid, --Secure, --Stable, --Tough (2).
  • Extraversion: --Reserved (7); Sociable, --Introverted, --Silent (6); Lively(5); Cheerful, --Passive, --Quiet, --Shy, --Withdrawn (4); Extraverted, Talkative, Vivacious, --Solitary, --Taciturn (3); Exuberant, Hyperactive, Merry, Open, Vigorous, --Boring, --Distant (2).
  • Openness: Original (5); Creative, Intellectual, Intelligent, Sharp (4); Clever, Gifted, Ironic, --Conservative, --Conventional (3); Artistic, Bright, Critical, Educated, Inventive, Receptive, Smart, Talented, Wise, Witty; --Backward, --Ignorant, --Incompetent, --Obedient, --Uneducated, --Unintelligent (2).
The -- means negative loading. The number in ()'s is how many studies mentioned those keywords out of a selection of them.

Emotionality is close to Big Five Neuroticism. H-H has some of Big Five Agreeableness.
 
Looking beyond the Big Five: A selective review of alternatives to the Big Five model of personality - ScienceDirect - mostly says that we ought to do research into additional personality factors, and cites a sizable number of them.
  • HEXACO - "Big Six"
  • Supernumerary personality traits - Conventionality, Seductiveness, Manipulativeness, Thriftiness, Humorousness, Integrity, Femininity, Religiosity, Risk-Taking, Egotism
  • Reduced versions- Machiavellian, Traditional, Masculine-Feminine
  • Psychobiological model (Cloninger et al.)
  • Dark Tetrad: Dark Triad (Machiavellianism, Psychopathy, Narcissism) & Everyday Sadism
  • Self-Defeating Personality Style
  • Emotional Intelligence

Higher-order factors of the Big Six – Similarities between Big Twos identified above the Big Five and the Big Six - ScienceDirect

The Big Two is alpha / beta or stability / plasticity.

Some studies find a Big One, a General Factor of Personality (GFP).

Big Five:
  • Alpha Agreeableness, Conscientiousness, (-) Neuroticism
  • Beta: Extraversion, Openness to Experience
  • GFP present: Alpha, Beta

Big Six:
  • Alpha Agreeableness, Conscientiousness, Honesty-Humility
  • Beta: Extraversion, Openness to Experience, (-) Emotionality
  • No GFP

Why does Neuroticism jump from Alpha to Beta when it becomes Emotionality?
 
These are certainly traits.

And they might appear stable if the person is in a stable non-threatening environment.

But they in no way explain behavior or could be used to predict behavior.

They are like skin color. A trait seen. But a trait that can change as an emotional response.
 
These are certainly traits.

And they might appear stable if the person is in a stable non-threatening environment.

But they in no way explain behavior or could be used to predict behavior.

They are like skin color. A trait seen. But a trait that can change as an emotional response.

I think they are most useful in helping people understand themselves more than anything else. I would not use them to hire someone, for example, or to try to predict behavior, as you said.
 
Conscientiousness: A Structural Assessment and Development of the Facets of Control Scales
Identifies three facets of Conscientiousness:
  • A "proactive aspect defined by Grit, Determination, Industriousness and Productiveness"
  • A "robust and consistent facet of Order and Organization"
  • An "inhibitory aspect of the trait was identified which was captured by Self-Control measures from the Chernyshenko Conscientiousness Scales, Prudence from the HEXACO and Impulse Control from the Facets of Control measure"
That is, diligence, orderliness, and self-control.

Under "Main Content": "This item is under embargo until July 20, 2022."
???

Seems like good work in homing in on the facet composition of C'ness. It would be interesting to see how other Big Five / HEXACO factors fare in this kind of analysis.


Integrating the HEXACO model with the Triarchic conceptualization of psychopathy - ScienceDirect
  • Boldness: -E +X
  • Meanness: -H +E
  • Disinhibition: -C
H = honesty-humility, E = emotionality (~ Big-5 neuroticism), X = eXtraversion, A = agreeableness, C = conscientiousness, O = openness
 
Is Hillary dishonest and Donald narcissistic? A HEXACO analysis of the presidential candidates' public personas - ScienceDirect
At ResearchGate: Is Hillary dishonest and Donald narcissistic? A HEXACO analysis of the presidential candidates' public personas - Visser20et20al.202016.pdf
As mentioned above, the HEXACO model offers several advantages over the Big Five model of personality including a plausible evolutionary basis (Lee & Ashton, 2012), replication in many languages and cultures (Ashton et al., 2006), and superior prediction of behavior, particularly deceitful, dishonest, and antisocial behavior (Lee & Ashton,2012), largely due to the inclusion of Honesty-Humility (de Vries et al., 2016). Further, in two large scale studies, the HEXACO has been shown to be an excellent predictor of particularly aversive personality types, namely psychopathy, narcissism, Machiavellianism, and sadism (Book et al., 2015; Book et al., 2016). More specifically, all of these personalities were predicted by low Honesty-Humility, Emotionality, Agreeableness, and Conscientiousness.


More generally, about HEXACO,
The HEXACO personality model (Ashton & Lee, 2007) proposes that there are six rather than five personality factors: Honesty-Humility (H), Emotionality (E), eXtraversion (X), Agreeableness (A), Conscientiousness (C), and Openness to Experience (O). A relatively new measure of personality, the HEXACO appears to have better theoretical, empirical, and cross-cultural support than the Big Five (de Vries, Tybur, Pollet, & van Vugt, 2016). In particular, the HEXACO delineates antisociality more clearly than the Big Five.

To begin with, people with low scores on Honesty-Humility are more likely to manipulate and exploit others, feel entitled and important, and are more likely to break rules for personal gain. There are 4 facets within the Honesty-Humility domain, namely: sincerity, fairness, greed avoidance, and modesty.

Low Emotionality scorers are emotionally detached and low on empathy, making them less likely to be concerned with the effect of their willing behavior on other people. They are also less likely to be worried in stressful situations, which may improve their crisis management skills. Facets within this domain include fearfulness, anxiety, sentimentality, and dependence.

High scores on eXtraversion are correlated with confidence, charisma, and sociability. The four facets in the domain of eXtraversion are social self-esteem, social boldness, sociability, and liveliness.

Agreeableness relates to the ability to forgive, being tolerant, and willing to compromise/cooperate with others. Low scorers tend to be vengeful, stubborn, and are more likely to react to provocation with anger. This domain is exemplified by facets measuring forgiveness, gentleness, flexibility, and patience.

People who are high on Conscientiousness are organized, disciplined, and make thoughtful decisions, whereas low scorers are impulsive in their decision making and are less concerned with quality of work. Conscientiousness is further split into facets measuring organization, diligence, perfectionism, and prudence.

The final subscale, Openness to Experience, is related to intellectual curiosity and the tendency to be interested in new or unconventional ideas. This domain includes 4 facets measuring aesthetic appreciation, inquisitiveness, creativity, and unconventionality.

There is also an interstitial scale directly measuring Altruism. People who score low on this scale are hard-hearted and less likely to engage in helping others.
 
PsyArXiv Preprints | Agreeableness in the HEXACO
We see little benefit to separating Honesty-Humility from the broader FFM Agreeableness domain. In our commentary, we summarize several studies showing that although lexically-based Big Five measures under-represent H-H content, the same cannot be said for FFM-based measures. We also indicate that contrary to claims by some advocates, FFM-based Agreeableness is more strongly related to the Dark Triad than H-H. Finally, we review a recent study examining the lower-order structure of FFM Agreeableness that failed to reveal a separate H-H factor, despite more than adequate representation of that content.
In other words, Big-5 Agreeableness includes HEXACO Honest-Humility along with HEXACO Agreeableness.


Uncovering the structure of agreeableness from self‐report measures - Crowe - 2018 - Journal of Personality - Wiley Online Library
"A five-factor solution consisting of facets labeled Compassion, Morality, Trust, Affability, and Modesty was identified as most appropriate"

Reopening Openness to Experience: A Network Analysis of Four Openness to Experience Inventories: Journal of Personality Assessment: Vol 101, No 6
Our results (N = 802) identified 10 distinct facets (variety-seeking, aesthetic appreciation, intellectual curiosity, diversity, openness to emotions, fantasy, imaginative, self-assessed intelligence, intellectual interests, and nontraditionalism) that largely replicate previous findings as well as three higher order aspects: two that are commonly found in the literature (intellect and experiencing; i.e., openness), and one novel aspect (open-mindedness).
From the paper,
  • Intellect
    • Intellectual interests -- Engagement in philosophy and discussing abstract, theoretical ideas
    • Self-assessed intelligence -- Perceived ability to think quickly, solve problems, and process information
    • Intellectual curiosity -- Enjoyment of learning new things, thinking about complexity, and reflecting on thoughts
  • Open-mindedness
    • Nontraditionalism -- Receptiveness to new ideas, cultures, and perspectives
    • Variety-seeking -- Willingness to explore new environments and try new ways of doing things
    • Diversity -- Embraces a variety of attitudes, beliefs, and lifestyles
  • Experiencing
    • Aesthetic appreciation -- Engagement in the arts and perceptual experiences
    • Openness to emotions -- Sensitivity to aesthetic emotions, complex feelings, and strong moods
    • Imaginative -- Ability to have original thoughts and a desire to create
    • Fantasy -- Tendency to daydream and mind wander

    I can't find much on the lower-order structure of extroversion or neuroticism.
 
PsyArXiv Preprints | Evaluating the Big Five as an Organizing Framework for Commonly Used Psychological Trait Scales
That meant recruiting a lot of subjects and giving them several psychological tests. One can then look for correlations between them.

Thus, worry has N +0.85, E -0.12, C +0.10, A +0.04, O 0.00

Behavioral self-inhibition has N +0.78, while self-compassion has N -0.68

Fun-seeking has E +0.56, C -0.27, A +0.14, O +0.13, N +0.04

Impulsivity (non-planning) has C -0.75, self-control C +0.70, grit (perseverance) has +0.65

Empathy (empathic conern) has A +0.76 and Aggression (verbal) has A -0.61

Need for cognition has O +0.61

Some of these scales rather obviously describe facets of the Big Five traits, but others don't seem to. Some traits are "interstitial", having contributions from more than one Big Five trait. Some others have variation outside of the Big Five. These "peripheral" scales usually had at least one of two features.

"First, many of these peripheral scales were comprised of items that began with “when” or “if,” indicating that some condition had to be fulfilled before the main clause of the item would be relevant"

"Second, many of these scale items included a common word (or synonym) that was either not in any other items or only rarely featured."
 
Looking back at early evolution, I found some more papers on dopamine and serotonin across the animal kingdom, in addition to earlier Ancestry of neuronal monoamine transporters in the Metazoa | Journal of Experimental Biology | The Company of Biologists

Frontiers | The Roles of Dopamine and Related Compounds in Reward-Seeking Behavior Across Animal Phyla | Behavioral Neuroscience
Motile animals actively seek out and gather resources they find rewarding, and this is an extremely powerful organizer and motivator of animal behavior. Mammalian studies have revealed interconnected neurobiological systems for reward learning, reward assessment, reinforcement and reward-seeking; all involving the biogenic amine dopamine. The neurobiology of reward-seeking behavioral systems is less well understood in invertebrates, but in many diverse invertebrate groups, reward learning and responses to food rewards also involve dopamine. The obvious exceptions are the arthropods in which the chemically related biogenic amine octopamine has a greater effect on reward learning and reinforcement than dopamine. Here we review the functions of these biogenic amines in behavioral responses to rewards in different animal groups, and discuss these findings in an evolutionary context.

IJMS | Free Full-Text | Serotonin in Animal Cognition and Behavior
Serotonin (5-hydroxytryptamine, 5-HT) is acknowledged as a major neuromodulator of nervous systems in both invertebrates and vertebrates. It has been proposed for several decades that it impacts animal cognition and behavior. In spite of a completely distinct organization of the 5-HT systems across the animal kingdom, several lines of evidence suggest that the influences of 5-HT on behavior and cognition are evolutionary conserved. In this review, we have selected some behaviors classically evoked when addressing the roles of 5-HT on nervous system functions. In particular, we focus on the motor activity, arousal, sleep and circadian rhythm, feeding, social interactions and aggressiveness, anxiety, mood, learning and memory, or impulsive/compulsive dimension and behavioral flexibility. The roles of 5-HT, illustrated in both invertebrates and vertebrates, show that it is more able to potentiate or mitigate the neuronal responses necessary for the fine-tuning of most behaviors, rather than to trigger or halt a specific behavior. 5-HT is, therefore, the prototypical neuromodulator fundamentally involved in the adaptation of all organisms across the animal kingdom.
So,
  • Dopamine -> reward systems -> Big Two plasticity
  • Serotonin -> modulation systems -> Big Two stability
both go back a long way, likely to the common ancestor of Bilateria and Cnidaria. So both mechanisms and their neurotransmitters are almost as old as nervous systems themselves.

The remaining animals with nervous systems are comb jellies, and there is the serious possibility that their nervous systems were the result of separate evolution.
 
More on different species:

  • Dog: Energy (Extraversion), Affection (Agreeableness), Emotional Reactivity (Neuroticism), Intelligence (Openness/Intellect). No counterpart of Conscientiousness, however.
  • Cat: Neuroticism, Extraversion, Dominance, Impulsiveness and Agreeableness.
  • Horse: Agreeableness, Extraversion, Neuroticism.
  • Parrot: Curiosity/neophilia (Extraversion), Anxiety/vigilance (Neuroticism)
Dominance and Impulsiveness seem related to Extraversion.
 
  • Old World simians (Catarrhini): (((human, chimp), gorilla), (rhesus monkey, vervet monkey))
  • Boreoeutheria: ((OWS, rat), ((dog, (hyena, cat)), (horse, donkey)))
  • Amniota: (Boreo, parrot)
  • Osteichthyes (Euteleostomi): (Amniota, guppy)
  • Eubilateria: (Oste, octopus)
Extraversion and neuroticism are found in all of them. Agreeableness was found among Boreoeutheria (northern-origin placental mammals).

I did a lot of searching for papers on personality variation in chickens, iguanas, and the like, but without much success. So finding papers on parrots was a big success -- they help fill the Boreo - fish gap.
 
More.

The performance of rooks in a cooperative task depends on their temperament | SpringerLink
In recent years, an increasing number of studies demonstrated the existence of consistent individual differences in behaviour, often referred to as differences in temperament or personality, in a wide range of animal species. There notably is a growing body of evidence showing that individuals differ in their propensity for risk taking or reacting to stressful situations.
Rook = Corvus frugilegus (crow-like bird) Roughly extraversion and neuroticism.

An experimental test of density‐dependent selection on temperament traits of activity, boldness and sociability - Le Galliard - 2015 - Journal of Evolutionary Biology - Wiley Online Library - tests on a common Eurasian lizard, Zootoca vivipara - activity, boldness and sociability

Dynamics of among-individual behavioral variation over adult lifespan in a wild insect | Behavioral Ecology | Oxford Academic - the cricket Gryllus campestris - tendency to leave a refuge, shyness, activity, and exploration

Personality variation in a clonal insect: The pea aphid, Acyrthosiphon pisum - Schuett - 2011 - Developmental Psychobiology - Wiley Online Library - "Here we show that individuals of clonal pea aphids exhibit consistent behavioral differences in their escape responses to a predator attack (dropping vs. nondropping off a plant)."

Polistes metricus queens exhibit personality variation and behavioral syndromes | Current Zoology | Oxford Academic - paper wasp - boldness, aggressiveness, exploration, and activity

Consistent crustaceans: the identification of stable behavioural syndromes in hermit crabs | SpringerLink - startle response, exploration and aggression

Most of these traits seem like versions of extraversion, but dropping/non-dropping and startle response seems like versions of neuroticism. As a comparison, Gosling et al. scored guppy fear-avoidance and octopus reactivity as neuroticism, and guppy approach and octopus boldness-avoidance as extraversion.
 
I found some personality research for a non-boreoeutherian mammal: Wild female African elephants (Loxodonta africana) exhibit personality traits of leadership and social integration. - PsycNET
Animal personalities have been demonstrated for almost 200 species, with stable dimensions of responses (aggressive to fearful; shy to bold) across contexts and with a heritable basis to these traits. As a long-lived and highly social species, elephants (Loxodonta africana) were expected to demonstrate complex dimensions to individual characteristics or personalities, which would be obvious to human observers and validated by behavioral observations. We used principal-components analysis of ratings on 26 behavioral adjectives applied to one social unit, coded as the EB family, which has been observed for 38 years. Eleven adult females were rated by four observers and found to have individually variable traits on four dimensions described by principal-components analysis. The first component was associated with effective and confident family leadership. Component 2 was age-related, and defined by playfulness, exploration and high levels of activity, suggesting both an experience and an age-related element to its structure. Component 3 represented gentleness and at its other extreme, aggression, and Component 4 was related to constancy (predictability and popularity), with both of these latter components reflecting social integration.
Evidence of extraversion, agreeableness, and likely openness.

A non-placental mammal: Individual traits influence vigilance in wild female eastern grey kangaroos | CSIRO PUBLISHING | Australian Journal of Zoology - neuroticism

Another reptile: Individual variation in boldness in turtles is consistent across assay conditions and behavioural measures in: Behaviour Volume 156 Issue 10 (2019) - extraversion

Another mollusk: Individual boldness is life stage-dependent and linked to dispersal in a hermaphrodite land snail | SpringerLink - extraversion
 
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